Animal physiology, immunology and developmental biology

This doctoral study programme is organized by the Faculty of Science in English and the studies are subject to tuition. There is an alternative option for the international applicants to be accepted in the free programme administered in Czech with the possibility of receiving a scholarship. The study language of the programme is still English (Czech is the administrative language). Before officially applying, please contact us at to find all the necessary information related to the scholarship and see our FAQ’s.

Submit an application

International applicants for doctoral study (Czech and Slovak Republics applicants not included)
Submission deadline until midnight 15 December 2022.

What will you learn?

The aim of the program is to prepare top specialists in specific scientific fields developed within the Department of Animal Physiology and Immunology at the Institute of Experimental Biology, Faculty of Science, Masaryk University. Graduates are prepared for further scientific careers at workplaces of basic or applied research at the international level, but also for possible employment in companies with a research background, in medical facilities and as professional lecturers.

The program focuses on the research formation of students in the rapidly developing fields of physiology, immunology and developmental biology. Students are trained to conduct research in the areas ranging from cellular to organ level to models of whole animal organisms from cells to human. Modern molecular biology techniques and state-of-the-art imaging techniques are applied. The program covers research topics ranging from veterinary through biomedical applications to general basic research issues. In particular, the most represented topic is the research of cellular signaling, which is crucial for understanding the processes controlling embryonic development, tissue differentiation and regeneration, as well as processes of carcinogenesis or immune regulation. The program integrates methodological approaches of molecular biology, molecular embryology and cell biology with morphological and behavioural approaches.

The course is designed to motivate prospective students through checkpoints. A greater number of methodologically oriented courses, internships and openness to contacts with biotechnology companies ensure interconnection with the practice of modern biological research. Emphasis is placed on using English as a basic tool for scientific work.

Students are motivated by the high professional quality of research in an open and friendly environment. The performance of individual students is appreciated both financially and, for example, through participation in prestigious international conferences.

Practical training

In accordance with international practice, the preferred form of dissertation is a collection of articles or manuscripts with an introductory article that logically links the individual articles and places them in the context of contemporary knowledge. In this initial article, the only author, which is a doctoral student, must prove his / her professional overview and orientation in the given field. The dissertation in this form usually contains three, but at least two articles or manuscripts of the author's articles. The best form is to submit the whole work in English.

Formal requirements for the work structure are given on the program website. The dissertation must be based on its own and original scientific results. A necessary condition for its acceptance is that its most important results are published or accepted for publication. The minimum requirements are met if the doctoral candidate is the first author of one work published in a journal of the first two quartiles of periodicals of the relevant focus (according to the Journal Citation Reports). Another article of the same quality (at least one) can be arbitrarily co-authored. If the doctoral dissertation articles are written with co-authors, the doctoral candidate in the dissertation must clearly define his / her specific contribution to each of them (not only the percentage). In cases of shared authoring, the Commission will generally apply stricter standards for publication quality. The work will be judged by 2-3 opponents, whereas at least two of them will have to be from another workplace, preferably foreign.

Career opportunities

Graduates of the doctoral study program are entitled to carry out independent scientific and research activities in the field of basic and applied research. In a broader sense, they are prepared to carry out all scientific work activities, and deal with conceptual issues, scientific-organizational activities, and teaching.

Admission requirements

Admission procedure
Admission board checks the candidates' professional capacities in an entrance interview that is conducted in English. The candidates submit a structured CV and in the course of the interview, they will present a brief proposal of their PhD project, will define the anticipated outputs, and suggest a methodical procedure. Written plan of candidate's suggested research project is also mandatory and will be submitted to the board. Recommended length of the plan is two pages and the structure is as follows: Title, Name of applicant, Name of supervisor, Home institution, Synopsis, Aims, Methodology, Financial covering. In a discussion with the admission board and in the presence of their supervisor they have to demonstrate that their respective thesis topic is scientifically well-grounded and experimentally manageable. A necessary prerequisite for admission is a demonstrable capability of professional conversation in English. The sufficient financial coverage of the candidate's scientific project will be assessed by the admission board within the terms of the entrance interview and further within the terms of the PhD Workshop in the 3rd semester.
The admission board assesses the candidates’ knowledge and prerequisites for the independent scientific work (0-200 points) and the capability of communication in English (0-100 points).

More information about admission process for international applicants in general can be found here.

Date of the entrance exam
The applicants will receive information about the entrance exam by e-mail usually at least 10 days before the exam.
Please, always check your e-mails, including spam folders.

Conditions of admission
To be admitted, the candidate must obtain at least 120 points in the professional and at least 60 points in the language part of the interview.
Successful applicants are informed of their acceptance by e-mail and subsequently receive an invitation to the enrolment.

Programme capacity
The capacity of a given programme is not fixed; students are admitted based on a decision by the Doctoral Board after assessing their aptitude for study and motivation.


2 May – 15 Dec 2022

Submit your application during this period

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Dissertation topics

Single-subject studies

Ecophysiology of photosynthesis and production in selected cryptogamic plants and tree species from Svalbard
Supervisor: prof. Ing. Miloš Barták, CSc.

Předmětem tématu doktorské disertační práce je sledování fotosyntetické aktivity vybraných druhů mechů, cévnatých rostlin (bylin) a dřevin pomocí biofyzikálních metod, zejména fluorescence chlorofylu in vivo a metod spektrální reflektance. Cílem doktorské disertační práce je kvantifikovat důležitost jednotlivých faktorů prostředí (chlad, mráz, nedostatek/přebytek vody, sluneční záření, teplota vzduchu, sněhová pokrývka) pro limitaci biofyzikálních a biochemických procesů fotosyntézy. V průběhu řešení bude uplatňován především přístup řízených laboratorních experimentů, součástí práce může být (v závislosti na úspěšnosti žádostí o granty) i terénní výzkum in situ.


prof. Ing. Miloš Barták, CSc.

Study information

Provided by Faculty of Science
Type of studies Doctoral
Mode full-time Yes
combined Yes
distance No
Study options single-subject studies Yes
single-subject studies with specialization No
major/minor studies No
Standard length of studies 4 years
Language of instruction English
Collaborating institutions
  • The Czech Academy of Sciences
  • Biofyzikální ústav AV ČR
Doctoral board and doctoral committees
Tuition fees
The studies are subject to tuition, fees are paid per academic year
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